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In the light-independent reactions, the chemical energy harvested during the light-dependent reactions drives the assembly of sugar molecules from carbon dioxide.
Therefore, although the light-independent reactions do not use light as a reactant, they require the products of the light-dependent reactions to function.
In addition, several enzymes of the light-independent reactions are activated by light.
The light-dependent reactions utilize certain molecules to temporarily store the energy: These are referred to as energy carriers.
The energy carriers that move energy from light-dependent reactions to light-independent reactions can be thought of as "full" because they are rich in energy.
After the energy is released, the "empty" energy carriers return to the light-dependent reaction to obtain more energy.
Moreover, the actual step that converts light energy into chemical energy takes place in a multiprotein complex called a photosystem, two types of which are found embedded in the thylakoid membrane: photosystem II (PSII) and photosystem I (PSI) .
The two complexes differ on the basis of what they oxidize (i.e., the source of the low-energy electron supply) and what they reduce (i.e., the place to which they deliver their energized electrons).
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Source: Boundless. “The Two Parts of Photosynthesis.” Boundless Biology. Boundless, 03 Jul. 2014. Retrieved 24 May. 2015 from https://www.boundless.com/biology/textbooks/boundless-biology-textbook/photosynthesis-8/overview-of-photosynthesis-80/the-two-parts-of-photosynthesis-373-11599/