An isotonic contraction where the muscle lengthens.
A muscle fiber generates tension through actin and myosin cross-bridge cycling. While under tension, the muscle may lengthen, shorten, or remain the same. Although the term contraction implies shortening, when referring to the muscular system, it means the generation of tension within a muscle fiber. Several types of muscle contractions occur and are defined by the changes in the length of the muscle during contraction.
Isotonic contractions maintain constant tension in the muscle as the muscle changes length. Isotonic muscle contractions can be either concentric or eccentric.
A concentric contraction is a type of muscle contraction in which the muscles shorten while generating force, overcoming resistance. For example, when lifting a heavy weight, a concentric contraction of the biceps would cause the arm to bend at the elbow, lifting the weight towards the shoulder. Cross-bridge cycling occurs, shortening the sarcomere, muscle fiber, and muscle.
An eccentric contraction results in the elongation of a muscle while the muscle is still generating force; in effect, resistance is greater than force generated. Eccentric contractions can be both voluntary and involuntary. For example, a voluntary eccentric contraction would be the controlled lowering of the heavy weight raised during the above concentric contraction. An involuntary eccentric contraction may occur when a weight is too great for a muscle to bear and so it is slowly lowered while under tension. Cross-bridge cycling occurs even though the sarcomere, muscle fiber, and muscle are lengthening, controlling the extension of the muscle.
In contrast to isotonic contractions, isometric contractions generate force without changing the length of the muscle, common in the muscles of the hand and forearm responsible for grip. Using the above example, the muscle contraction required to grip but not move a heavy object prior to lifting would be isometric. Isometric contractions are frequently used to maintain posture.
Isometric contractions are sometimes described as yielding or overcoming.
A yielding contraction occurs when a muscle contraction is opposed by resistance. For example, when holding a heavy weight steady, neither raising nor lowering it.
An overcoming contraction occurs when a muscle contraction is opposed by an immovable object, such as the contraction generated in the muscles when pushing against a wall.
In both instances, cross-bridge cycling is maintaining tension in the muscle; the sarcomere, muscle fibers, and muscle are not changing length.